1. Compared to other presbyornithids, Willaru appeared to have been specialised to a more terrestrial lifestyle, based on its tarsometatarsal morphology.

2. Arthrodesis of the tarsometatarsal (TMT) joint is performed for treatment of arthritis of variable extent with or without deformity, in the setting of idiopathic osteoarthritis or posttraumatic or inflammatory arthritis, and for correction of neuropathic deformity.